494 BC

Having successfully captured several of the revolting Greek city-states, the Persians under Artaphernes lay siege to Miletus. The decisive Battle of Lade is fought at the island of Lade, near Miletus' port. Although out-numbered, the Greek fleet appears to be winning the battle until the ships from Samos and Lesbos retreat. The sudden defection turns the tide of battle, and the remaining Greek fleet is completely destroyed. Miletus surrenders shortly thereafter, and the Ionian Revolt comes to an end.